The Devonian is a geologic period and system of the Paleozoic Era spanning from the end of the Silurian Period, about  Mya (million years ago), to the beginning of the Carboniferous Period, about . It is named after Devon, England, where rocks from this period were first studied. The Devonian period experienced the first significant adaptive radiation of terrestrial life. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. Various terrestrial arthropods also became well-established. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the "Age of Fish". The first ray-finned and lobe-finned bony fish appeared, while the placoderms began dominating almost every known aquatic environment.