An
evolutionary landscape is a metaphor, a construct used to think about and visualize the processes of
evolution (e.g.
natural selection and
genetic drift) acting on a biological entity (e.g., a
gene,
protein,
population,
species). This entity can be viewed as searching or moving through a
search space. For example, the search space of a gene would be all possible nucleotide sequences. The search space is only part of an evolutionary landscape. The final component is the "y-axis," which is usually
fitness. Each value along the search space can result in a high or low fitness for the entity. If small movements through search space cause changes in fitness that are relatively small, then the landscape is considered smooth. Smooth landscapes happen when most fixed mutations have little to no effect on fitness, which is what one would expect with
the neutral theory of molecular evolution. In contrast, if small movements result in large changes in fitness, then the landscape is said to be rugged. In either case, movement tends to be toward areas of higher fitness, though usually not the global optima.